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# POD documentation - main docs before the code |
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=head1 NAME |
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5
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Module which implements a newick string parser as a finite state |
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machine which enables it to parse the full Newick specification. |
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Taken largely from the Ensembl Compara file with the same name |
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(Bio::EnsEMBL::Compara::Graph::NewickParser), this module adapts the |
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parser to work with BioPerl's event handler-based parsing scheme. |
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This module is used by nhx.pm and newick.pm, and is NOT called |
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directly. Instead, both of those parsing modules extend this module in |
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order to gain access to the main parsing method. |
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=head1 SYNOPSIS |
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# From newick.pm |
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use base qw(Bio::TreeIO Bio::TreeIO::NewickParser); |
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# in the next_tree method... |
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$self->parse_newick($_); |
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=head1 DESCRIPTION |
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This module correctly parses the Newick and NHX formats, sending calls |
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to the BioPerl TreeEventHandler when appropriate in order to build and |
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populate the node objects. |
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=head1 FEEDBACK |
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=head2 Mailing Lists |
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User feedback is an integral part of the evolution of this and other |
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Bioperl modules. Send your comments and suggestions preferably to the |
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Bioperl mailing list. Your participation is much appreciated. |
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bioperl-l@bioperl.org - General discussion |
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http://bioperl.org/wiki/Mailing_lists - About the mailing lists |
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41
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=head2 Support |
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43
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Please direct usage questions or support issues to the mailing list: |
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45
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I |
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rather than to the module maintainer directly. Many experienced and |
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reponsive experts will be able look at the problem and quickly |
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address it. Please include a thorough description of the problem |
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with code and data examples if at all possible. |
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51
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52
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=head2 Reporting Bugs |
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53
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54
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Report bugs to the Bioperl bug tracking system to help us keep track |
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55
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of the bugs and their resolution. Bug reports can be submitted via the |
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56
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web: |
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58
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https://github.com/bioperl/bioperl-live/issues |
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59
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60
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=head1 AUTHOR - Jessica Severin (EnsEMBL implementation), Greg Jordan (BioPerl adaptation) |
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61
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62
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=cut |
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63
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64
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package Bio::TreeIO::NewickParser; |
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65
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66
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15
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15
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100
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use strict; |
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15
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52
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15
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403
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67
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15
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59
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use base qw(Bio::Root::Root); |
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15
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26
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15
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13728
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68
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69
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sub parse_newick { |
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70
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126
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126
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0
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226
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my $self = shift; |
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71
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126
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271
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my $newick = shift; |
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72
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73
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126
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100
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477
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$newick = $newick . ";" unless ($newick =~ m/;/); |
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74
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75
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126
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202
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my $count=1; |
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76
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126
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355
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my $debug = $self->verbose; |
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77
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126
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374
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my $token = next_token(\$newick, "(;"); |
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78
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126
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227
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my $state=1; |
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79
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126
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195
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my $bracket_level = 0; |
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80
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81
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126
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477
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$self->_start('tree'); |
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82
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83
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126
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297
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my $leaf_flag = 0; |
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84
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85
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126
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293
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while(defined($token)) { |
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86
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# backwards-compat. with 5.8.1, no Switch (but we hate if-elsif-ad-infinitum |
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87
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23135
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100
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46032
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if ($state == 1) { #new node |
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100
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100
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100
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50
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0
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88
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89
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4627
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9627
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$self->_start('node'); |
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90
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91
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4627
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16093
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$self->debug(" -> [$token]\n"); |
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92
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4627
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100
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9194
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if($token eq '(') { #create new set |
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93
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2223
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50
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3910
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$self->debug(" create set\n") if($debug); |
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94
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2223
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3962
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$token = next_token(\$newick, "[(:,)"); |
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95
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2223
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3034
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$state = 1; |
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96
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2223
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4050
|
$bracket_level++; |
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97
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} else { |
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98
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2404
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3190
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$state = 2; |
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99
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2404
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3742
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$leaf_flag = 1; |
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100
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} |
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101
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} elsif ($state == 2) { #naming a node |
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102
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4627
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100
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13625
|
if(!($token =~ /[\[\:\,\)\;]/)) { |
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103
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104
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2488
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100
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100
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5659
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if (!$leaf_flag && $self->param('internal_node_id') eq 'bootstrap') { |
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105
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4
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17
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$self->_start('bootstrap'); |
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106
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4
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18
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$self->_chars($token); |
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107
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4
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16
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$self->_end('bootstrap'); |
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108
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4
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12
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$token = ''; |
|
109
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} |
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110
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111
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2488
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5074
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$self->_start('id'); |
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112
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2488
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6627
|
$self->_chars($token); |
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113
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2488
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6280
|
$self->_end('id'); |
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114
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115
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2488
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50
|
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5761
|
$self->debug(" naming leaf\n") if ($debug); |
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116
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2488
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5056
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$token = next_token(\$newick, "[:,);"); |
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117
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} |
|
118
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4627
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7801
|
$state = 3; |
|
119
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} elsif ($state == 3) { # optional : and distance |
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120
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4627
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100
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7411
|
if($token eq ':') { |
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100
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121
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4131
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6577
|
$token = next_token(\$newick, "[,);"); |
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122
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123
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4131
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9364
|
$self->_start('branch_length'); |
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124
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4131
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10771
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$self->_chars($token); |
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125
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4131
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9277
|
$self->_end('branch_length'); |
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126
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127
|
4131
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10013
|
$token = next_token(\$newick, ",);"); #move to , or ) |
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128
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} elsif ($token eq '[') { # NHX tag without previous blength |
|
129
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15
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42
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$token .= next_token(\$newick, ",);"); |
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130
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} |
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131
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4627
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8424
|
$state = 4; |
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132
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} elsif ($state == 4) { # optional NHX tags |
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133
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4627
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100
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12335
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if($token =~ /\[\&\&NHX/) { |
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100
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134
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# careful: this regexp gets rid of all NHX wrapping in one step |
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135
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136
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478
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1093
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$self->_start('nhx_tag'); |
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137
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478
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2349
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$token =~ /\[\&\&NHX\:(\S+)\]/; |
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138
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478
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50
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1721
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if ($1) { |
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139
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# NHX may be empty, presumably at end of file, just before ";" |
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140
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478
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1286
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my @attributes = split ':', $1; |
|
141
|
478
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880
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foreach my $attribute (@attributes) { |
|
142
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778
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1464
|
$attribute =~ s/\s+//; |
|
143
|
778
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2007
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my($key,$value) = split '=', $attribute; |
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144
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145
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778
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1894
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$self->_start('tag_name'); |
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146
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778
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1860
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$self->_chars($key); |
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147
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778
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1716
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$self->_end('tag_name'); |
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148
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149
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778
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2005
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$self->_start('tag_value'); |
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150
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778
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2130
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$self->_chars($value); |
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151
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778
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1531
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$self->_end('tag_value'); |
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152
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} |
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153
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} |
|
154
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478
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1156
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$self->_end('nhx_tag'); |
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155
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156
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478
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1221
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$token = next_token(\$newick, ",);"); #move to , or ) |
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157
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} elsif ($token =~ /\[/) { |
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158
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# This is a hack to make AMPHORA2 work |
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159
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4
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50
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|
29
|
if ($token =~ /\[(\S+)\]/) { |
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160
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4
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18
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$self->_start('bootstrap'); |
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161
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4
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19
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$self->_chars($1); |
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162
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4
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16
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$self->_end('bootstrap'); |
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163
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} |
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164
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4
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20
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$token = next_token(\$newick, ",);"); } |
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165
|
4627
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|
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7593
|
$state = 5; |
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166
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} elsif ($state == 5) { # end node |
|
167
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4627
|
100
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|
8770
|
if($token eq ')') { |
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100
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50
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168
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169
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2223
|
|
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|
5013
|
$self->_end('node'); |
|
170
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|
171
|
2223
|
|
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|
5769
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$token = next_token(\$newick, "[:,);"); |
|
172
|
2223
|
100
|
66
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|
7958
|
if (defined $token && $token eq '[') { |
|
173
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# It is possible to have anonymous internal nodes w/ no name |
|
174
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# and no blength but with NHX tags |
|
175
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# |
|
176
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# We use leaf_flag=0 to let the parser know that it's labeling an internal |
|
177
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|
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# node. This affects how potential bootstrap values are handled in state 2. |
|
178
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8
|
|
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|
15
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$leaf_flag = 0; |
|
179
|
8
|
|
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|
16
|
$state = 2; |
|
180
|
|
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|
|
} else { |
|
181
|
2215
|
|
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|
|
2718
|
$leaf_flag = 0; |
|
182
|
2215
|
|
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|
|
2532
|
$state = 2; |
|
183
|
|
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} |
|
184
|
2223
|
|
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|
4038
|
$bracket_level--; |
|
185
|
|
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|
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} elsif($token eq ',') { |
|
186
|
|
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187
|
2278
|
|
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|
4966
|
$self->_end('node'); |
|
188
|
|
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189
|
2278
|
|
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|
|
6565
|
$token = next_token(\$newick, "[(:,)"); #can be un_blengthed nhx nodes |
|
190
|
2278
|
|
|
|
|
4724
|
$state=1; |
|
191
|
|
|
|
|
|
|
} elsif($token eq ';') { |
|
192
|
|
|
|
|
|
|
#done with tree |
|
193
|
126
|
50
|
|
|
|
336
|
$self->throw("parse error: unbalanced ()\n") if($bracket_level ne 0); |
|
194
|
|
|
|
|
|
|
|
|
195
|
126
|
|
|
|
|
324
|
$self->_end('node'); |
|
196
|
126
|
|
|
|
|
404
|
$self->_end('tree'); |
|
197
|
|
|
|
|
|
|
|
|
198
|
126
|
|
|
|
|
399
|
$token = next_token(\$newick, "("); |
|
199
|
126
|
|
|
|
|
380
|
$state=13; |
|
200
|
|
|
|
|
|
|
} else { |
|
201
|
0
|
|
|
|
|
0
|
$self->debug("[$token]]\n"); |
|
202
|
0
|
|
|
|
|
0
|
$self->throw("parse error: expected ; or ) or ,\n"); |
|
203
|
|
|
|
|
|
|
} |
|
204
|
|
|
|
|
|
|
} elsif ($state == 13) { |
|
205
|
0
|
|
|
|
|
0
|
$self->throw("parse error: nothing expected after ;"); |
|
206
|
|
|
|
|
|
|
} |
|
207
|
|
|
|
|
|
|
} |
|
208
|
|
|
|
|
|
|
|
|
209
|
126
|
50
|
|
|
|
350
|
if ($self->_eventHandler->within_element('tree')) { |
|
210
|
0
|
|
|
|
|
0
|
$self->_end('node'); |
|
211
|
0
|
|
|
|
|
0
|
$self->_end('tree'); |
|
212
|
|
|
|
|
|
|
} |
|
213
|
|
|
|
|
|
|
} |
|
214
|
|
|
|
|
|
|
|
|
215
|
|
|
|
|
|
|
sub _chars { |
|
216
|
8183
|
|
|
8183
|
|
9370
|
my $self = shift; |
|
217
|
8183
|
|
|
|
|
9822
|
my $chars = shift; |
|
218
|
|
|
|
|
|
|
|
|
219
|
8183
|
|
|
|
|
12740
|
$self->_eventHandler->characters($chars); |
|
220
|
|
|
|
|
|
|
} |
|
221
|
|
|
|
|
|
|
|
|
222
|
|
|
|
|
|
|
sub _start { |
|
223
|
13414
|
|
|
13414
|
|
14552
|
my $self = shift; |
|
224
|
13414
|
|
|
|
|
14386
|
my $name = shift; |
|
225
|
|
|
|
|
|
|
|
|
226
|
13414
|
|
|
|
|
24096
|
$self->_eventHandler->start_element({Name=>$name}); |
|
227
|
|
|
|
|
|
|
} |
|
228
|
|
|
|
|
|
|
|
|
229
|
|
|
|
|
|
|
sub _end { |
|
230
|
13414
|
|
|
13414
|
|
15297
|
my $self = shift; |
|
231
|
13414
|
|
|
|
|
14965
|
my $name = shift; |
|
232
|
|
|
|
|
|
|
|
|
233
|
13414
|
|
|
|
|
22714
|
$self->_eventHandler->end_element({Name=>$name}); |
|
234
|
|
|
|
|
|
|
} |
|
235
|
|
|
|
|
|
|
|
|
236
|
|
|
|
|
|
|
sub next_token { |
|
237
|
18223
|
|
|
18223
|
0
|
20365
|
my $string = shift; |
|
238
|
18223
|
|
|
|
|
19456
|
my $delim = shift; |
|
239
|
|
|
|
|
|
|
|
|
240
|
18223
|
|
|
|
|
41446
|
$$string =~ s/^(\s)+//; |
|
241
|
|
|
|
|
|
|
|
|
242
|
18223
|
100
|
|
|
|
29833
|
return undef unless(length($$string)); |
|
243
|
|
|
|
|
|
|
|
|
244
|
|
|
|
|
|
|
#print("input =>$$string\n"); |
|
245
|
|
|
|
|
|
|
#print("delim =>$delim\n"); |
|
246
|
18097
|
|
|
|
|
19554
|
my $index=undef; |
|
247
|
|
|
|
|
|
|
|
|
248
|
18097
|
|
|
|
|
69022
|
my @delims = split(/ */, $delim); |
|
249
|
18097
|
|
|
|
|
29636
|
foreach my $dl (@delims) { |
|
250
|
76720
|
|
|
|
|
89838
|
my $pos = index($$string, $dl); |
|
251
|
76720
|
100
|
|
|
|
99677
|
if($pos>=0) { |
|
252
|
62666
|
100
|
|
|
|
80420
|
$index = $pos unless(defined($index)); |
|
253
|
62666
|
100
|
|
|
|
91836
|
$index = $pos if($pos<$index); |
|
254
|
|
|
|
|
|
|
} |
|
255
|
|
|
|
|
|
|
} |
|
256
|
18097
|
50
|
|
|
|
26439
|
unless(defined($index)) { |
|
257
|
|
|
|
|
|
|
# have to call as class here (this is not an instance method) |
|
258
|
0
|
|
|
|
|
0
|
Bio::Root::Root->throw("couldn't find delimiter $delim\n $$string"); |
|
259
|
|
|
|
|
|
|
} |
|
260
|
|
|
|
|
|
|
|
|
261
|
18097
|
|
|
|
|
22175
|
my $token =''; |
|
262
|
|
|
|
|
|
|
|
|
263
|
18097
|
100
|
|
|
|
25526
|
if($index==0) { |
|
264
|
10996
|
|
|
|
|
17218
|
$token = substr($$string,0,1); |
|
265
|
10996
|
|
|
|
|
21290
|
$$string = substr($$string, 1); |
|
266
|
|
|
|
|
|
|
} else { |
|
267
|
7101
|
|
|
|
|
10916
|
$token = substr($$string, 0, $index); |
|
268
|
7101
|
|
|
|
|
13030
|
$$string = substr($$string, $index); |
|
269
|
|
|
|
|
|
|
} |
|
270
|
|
|
|
|
|
|
|
|
271
|
|
|
|
|
|
|
#print(" token =>$token\n"); |
|
272
|
|
|
|
|
|
|
#print(" outstring =>$$string\n\n"); |
|
273
|
|
|
|
|
|
|
|
|
274
|
18097
|
|
|
|
|
39266
|
return $token; |
|
275
|
|
|
|
|
|
|
} |
|
276
|
|
|
|
|
|
|
|
|
277
|
|
|
|
|
|
|
|
|
278
|
|
|
|
|
|
|
|
|
279
|
|
|
|
|
|
|
1; |