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# POD documentation - main docs before the code |
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=head1 NAME |
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Module which implements a newick string parser as a finite state |
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machine which enables it to parse the full Newick specification. |
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Taken largely from the Ensembl Compara file with the same name |
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(Bio::EnsEMBL::Compara::Graph::NewickParser), this module adapts the |
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parser to work with BioPerl's event handler-based parsing scheme. |
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This module is used by nhx.pm and newick.pm, and is NOT called |
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directly. Instead, both of those parsing modules extend this module in |
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order to gain access to the main parsing method. |
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=head1 SYNOPSIS |
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# From newick.pm |
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use base qw(Bio::TreeIO Bio::TreeIO::NewickParser); |
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# in the next_tree method... |
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$self->parse_newick($_); |
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=head1 DESCRIPTION |
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This module correctly parses the Newick and NHX formats, sending calls |
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to the BioPerl TreeEventHandler when appropriate in order to build and |
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populate the node objects. |
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=head1 FEEDBACK |
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=head2 Mailing Lists |
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User feedback is an integral part of the evolution of this and other |
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Bioperl modules. Send your comments and suggestions preferably to the |
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Bioperl mailing list. Your participation is much appreciated. |
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bioperl-l@bioperl.org - General discussion |
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http://bioperl.org/wiki/Mailing_lists - About the mailing lists |
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=head2 Support |
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Please direct usage questions or support issues to the mailing list: |
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I |
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rather than to the module maintainer directly. Many experienced and |
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reponsive experts will be able look at the problem and quickly |
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address it. Please include a thorough description of the problem |
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with code and data examples if at all possible. |
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=head2 Reporting Bugs |
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54
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Report bugs to the Bioperl bug tracking system to help us keep track |
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of the bugs and their resolution. Bug reports can be submitted via the |
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web: |
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58
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https://github.com/bioperl/bioperl-live/issues |
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60
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=head1 AUTHOR - Jessica Severin (EnsEMBL implementation), Greg Jordan (BioPerl adaptation) |
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=cut |
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64
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package Bio::TreeIO::NewickParser; |
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66
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use strict; |
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17
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16
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432
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67
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46
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use base qw(Bio::Root::Root); |
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16
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16
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16
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13462
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68
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69
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sub parse_newick { |
70
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153
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153
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0
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165
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my $self = shift; |
71
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153
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170
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my $newick = shift; |
72
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73
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153
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100
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450
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$newick = $newick . ";" unless ($newick =~ m/;/); |
74
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75
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153
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134
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my $count=1; |
76
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153
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321
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my $debug = $self->verbose; |
77
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153
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300
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my $token = next_token(\$newick, "(;"); |
78
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153
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169
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my $state=1; |
79
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153
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136
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my $bracket_level = 0; |
80
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81
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153
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366
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$self->_start('tree'); |
82
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83
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153
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243
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my $leaf_flag = 0; |
84
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85
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153
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307
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while(defined($token)) { |
86
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# backwards-compat. with 5.8.1, no Switch (but we hate if-elsif-ad-infinitum |
87
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24255
|
100
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41660
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if ($state == 1) { #new node |
|
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100
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100
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100
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50
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0
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88
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89
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4851
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6073
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$self->_start('node'); |
90
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91
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4851
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10349
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$self->debug(" -> [$token]\n"); |
92
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4851
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100
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6752
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if($token eq '(') { #create new set |
93
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2308
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50
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2782
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$self->debug(" create set\n") if($debug); |
94
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2308
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2647
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$token = next_token(\$newick, "[(:,)"); |
95
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2308
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1708
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$state = 1; |
96
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2308
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3274
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$bracket_level++; |
97
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} else { |
98
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2543
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1934
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$state = 2; |
99
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2543
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3509
|
$leaf_flag = 1; |
100
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} |
101
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} elsif ($state == 2) { #naming a node |
102
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4851
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100
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10052
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if(!($token =~ /[\[\:\,\)\;]/)) { |
103
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104
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2632
|
100
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100
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4422
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if (!$leaf_flag && $self->param('internal_node_id') eq 'bootstrap') { |
105
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4
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8
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$self->_start('bootstrap'); |
106
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4
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7
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$self->_chars($token); |
107
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4
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8
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$self->_end('bootstrap'); |
108
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4
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7
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$token = ''; |
109
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} |
110
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111
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2632
|
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3149
|
$self->_start('id'); |
112
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2632
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4078
|
$self->_chars($token); |
113
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2632
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3778
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$self->_end('id'); |
114
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115
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2632
|
50
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4481
|
$self->debug(" naming leaf\n") if ($debug); |
116
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2632
|
|
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3220
|
$token = next_token(\$newick, "[:,);"); |
117
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} |
118
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4851
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6780
|
$state = 3; |
119
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} elsif ($state == 3) { # optional : and distance |
120
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4851
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100
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5936
|
if($token eq ':') { |
|
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100
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121
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4325
|
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4926
|
$token = next_token(\$newick, "[,);"); |
122
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123
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4325
|
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6325
|
$self->_start('branch_length'); |
124
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4325
|
|
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6761
|
$self->_chars($token); |
125
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4325
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5936
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$self->_end('branch_length'); |
126
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127
|
4325
|
|
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7084
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$token = next_token(\$newick, ",);"); #move to , or ) |
128
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} elsif ($token eq '[') { # NHX tag without previous blength |
129
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15
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25
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$token .= next_token(\$newick, ",);"); |
130
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} |
131
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4851
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7703
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$state = 4; |
132
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} elsif ($state == 4) { # optional NHX tags |
133
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4851
|
100
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10246
|
if($token =~ /\[\&\&NHX/) { |
|
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100
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134
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# careful: this regexp gets rid of all NHX wrapping in one step |
135
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136
|
478
|
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643
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$self->_start('nhx_tag'); |
137
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478
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|
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1426
|
$token =~ /\[\&\&NHX\:(\S+)\]/; |
138
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478
|
50
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1182
|
if ($1) { |
139
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# NHX may be empty, presumably at end of file, just before ";" |
140
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478
|
|
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833
|
my @attributes = split ':', $1; |
141
|
478
|
|
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490
|
foreach my $attribute (@attributes) { |
142
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778
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|
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904
|
$attribute =~ s/\s+//; |
143
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778
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|
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1382
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my($key,$value) = split '=', $attribute; |
144
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145
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778
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1158
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$self->_start('tag_name'); |
146
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778
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1153
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$self->_chars($key); |
147
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778
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1017
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$self->_end('tag_name'); |
148
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149
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778
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1118
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$self->_start('tag_value'); |
150
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778
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1051
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$self->_chars($value); |
151
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778
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1042
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$self->_end('tag_value'); |
152
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} |
153
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} |
154
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478
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585
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$self->_end('nhx_tag'); |
155
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156
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478
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732
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$token = next_token(\$newick, ",);"); #move to , or ) |
157
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} elsif ($token =~ /\[/) { |
158
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# This is a hack to make AMPHORA2 work |
159
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4
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50
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10
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if ($token =~ /\[(\S+)\]/) { |
160
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4
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8
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$self->_start('bootstrap'); |
161
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4
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5
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$self->_chars($1); |
162
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4
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104
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$self->_end('bootstrap'); |
163
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} |
164
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4
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9
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$token = next_token(\$newick, ",);"); } |
165
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4851
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6045
|
$state = 5; |
166
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} elsif ($state == 5) { # end node |
167
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4851
|
100
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6775
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if($token eq ')') { |
|
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100
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50
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168
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169
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2308
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|
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3053
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$self->_end('node'); |
170
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171
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2308
|
|
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|
3901
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$token = next_token(\$newick, "[:,);"); |
172
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2308
|
100
|
66
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|
7051
|
if (defined $token && $token eq '[') { |
173
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|
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|
# It is possible to have anonymous internal nodes w/ no name |
174
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|
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# and no blength but with NHX tags |
175
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# |
176
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# We use leaf_flag=0 to let the parser know that it's labeling an internal |
177
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# node. This affects how potential bootstrap values are handled in state 2. |
178
|
8
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|
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|
|
13
|
$leaf_flag = 0; |
179
|
8
|
|
|
|
|
12
|
$state = 2; |
180
|
|
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|
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} else { |
181
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2300
|
|
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1743
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$leaf_flag = 0; |
182
|
2300
|
|
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|
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1648
|
$state = 2; |
183
|
|
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|
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|
|
} |
184
|
2308
|
|
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|
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3608
|
$bracket_level--; |
185
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|
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|
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|
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} elsif($token eq ',') { |
186
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|
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187
|
2390
|
|
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|
|
3064
|
$self->_end('node'); |
188
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|
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|
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189
|
2390
|
|
|
|
|
4110
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$token = next_token(\$newick, "[(:,)"); #can be un_blengthed nhx nodes |
190
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2390
|
|
|
|
|
4293
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$state=1; |
191
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|
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} elsif($token eq ';') { |
192
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|
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#done with tree |
193
|
153
|
50
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|
340
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$self->throw("parse error: unbalanced ()\n") if($bracket_level ne 0); |
194
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|
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195
|
153
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|
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282
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$self->_end('node'); |
196
|
153
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|
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|
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341
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$self->_end('tree'); |
197
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|
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198
|
153
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|
306
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$token = next_token(\$newick, "("); |
199
|
153
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|
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360
|
$state=13; |
200
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|
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} else { |
201
|
0
|
|
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|
|
0
|
$self->debug("[$token]]\n"); |
202
|
0
|
|
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|
|
0
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$self->throw("parse error: expected ; or ) or ,\n"); |
203
|
|
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} |
204
|
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} elsif ($state == 13) { |
205
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0
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|
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|
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0
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$self->throw("parse error: nothing expected after ;"); |
206
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|
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|
} |
207
|
|
|
|
|
|
|
} |
208
|
|
|
|
|
|
|
|
209
|
153
|
50
|
|
|
|
278
|
if ($self->_eventHandler->within_element('tree')) { |
210
|
0
|
|
|
|
|
0
|
$self->_end('node'); |
211
|
0
|
|
|
|
|
0
|
$self->_end('tree'); |
212
|
|
|
|
|
|
|
} |
213
|
|
|
|
|
|
|
} |
214
|
|
|
|
|
|
|
|
215
|
|
|
|
|
|
|
sub _chars { |
216
|
8521
|
|
|
8521
|
|
6293
|
my $self = shift; |
217
|
8521
|
|
|
|
|
6229
|
my $chars = shift; |
218
|
|
|
|
|
|
|
|
219
|
8521
|
|
|
|
|
10160
|
$self->_eventHandler->characters($chars); |
220
|
|
|
|
|
|
|
} |
221
|
|
|
|
|
|
|
|
222
|
|
|
|
|
|
|
sub _start { |
223
|
14003
|
|
|
14003
|
|
10043
|
my $self = shift; |
224
|
14003
|
|
|
|
|
9751
|
my $name = shift; |
225
|
|
|
|
|
|
|
|
226
|
14003
|
|
|
|
|
18319
|
$self->_eventHandler->start_element({Name=>$name}); |
227
|
|
|
|
|
|
|
} |
228
|
|
|
|
|
|
|
|
229
|
|
|
|
|
|
|
sub _end { |
230
|
14003
|
|
|
14003
|
|
10515
|
my $self = shift; |
231
|
14003
|
|
|
|
|
9987
|
my $name = shift; |
232
|
|
|
|
|
|
|
|
233
|
14003
|
|
|
|
|
16857
|
$self->_eventHandler->end_element({Name=>$name}); |
234
|
|
|
|
|
|
|
} |
235
|
|
|
|
|
|
|
|
236
|
|
|
|
|
|
|
sub next_token { |
237
|
19091
|
|
|
19091
|
0
|
13933
|
my $string = shift; |
238
|
19091
|
|
|
|
|
13058
|
my $delim = shift; |
239
|
|
|
|
|
|
|
|
240
|
19091
|
|
|
|
|
28079
|
$$string =~ s/^(\s)+//; |
241
|
|
|
|
|
|
|
|
242
|
19091
|
100
|
|
|
|
24817
|
return undef unless(length($$string)); |
243
|
|
|
|
|
|
|
|
244
|
|
|
|
|
|
|
#print("input =>$$string\n"); |
245
|
|
|
|
|
|
|
#print("delim =>$delim\n"); |
246
|
18938
|
|
|
|
|
13126
|
my $index=undef; |
247
|
|
|
|
|
|
|
|
248
|
18938
|
|
|
|
|
54844
|
my @delims = split(/ */, $delim); |
249
|
18938
|
|
|
|
|
19797
|
foreach my $dl (@delims) { |
250
|
80262
|
|
|
|
|
72259
|
my $pos = index($$string, $dl); |
251
|
80262
|
100
|
|
|
|
88791
|
if($pos>=0) { |
252
|
65273
|
100
|
|
|
|
71293
|
$index = $pos unless(defined($index)); |
253
|
65273
|
100
|
|
|
|
81828
|
$index = $pos if($pos<$index); |
254
|
|
|
|
|
|
|
} |
255
|
|
|
|
|
|
|
} |
256
|
18938
|
50
|
|
|
|
21834
|
unless(defined($index)) { |
257
|
|
|
|
|
|
|
# have to call as class here (this is not an instance method) |
258
|
0
|
|
|
|
|
0
|
Bio::Root::Root->throw("couldn't find delimiter $delim\n $$string"); |
259
|
|
|
|
|
|
|
} |
260
|
|
|
|
|
|
|
|
261
|
18938
|
|
|
|
|
14917
|
my $token =''; |
262
|
|
|
|
|
|
|
|
263
|
18938
|
100
|
|
|
|
20595
|
if($index==0) { |
264
|
11499
|
|
|
|
|
11963
|
$token = substr($$string,0,1); |
265
|
11499
|
|
|
|
|
14827
|
$$string = substr($$string, 1); |
266
|
|
|
|
|
|
|
} else { |
267
|
7439
|
|
|
|
|
7498
|
$token = substr($$string, 0, $index); |
268
|
7439
|
|
|
|
|
9153
|
$$string = substr($$string, $index); |
269
|
|
|
|
|
|
|
} |
270
|
|
|
|
|
|
|
|
271
|
|
|
|
|
|
|
#print(" token =>$token\n"); |
272
|
|
|
|
|
|
|
#print(" outstring =>$$string\n\n"); |
273
|
|
|
|
|
|
|
|
274
|
18938
|
|
|
|
|
29568
|
return $token; |
275
|
|
|
|
|
|
|
} |
276
|
|
|
|
|
|
|
|
277
|
|
|
|
|
|
|
|
278
|
|
|
|
|
|
|
|
279
|
|
|
|
|
|
|
1; |